Rabidosa rabida

Ge­o­graphic Range

Ra­bidosa ra­bida is very com­mon and widely dis­trib­uted. It is often recorded as a wan­der­ing spi­der. It is typ­i­cally found in east to cen­tral Texas and Ok­la­homa, north­ward to Ne­braska. It has fre­quently been traced as far east as Maine and as far south as Florida. (Jack­man, 1997)

Habi­tat

Ra­bidosa ra­bida is­most often found in wooded areas, cot­ton fields, and build­ings. These spi­ders usu­ally re­side among lit­ter, rub­bish, in holes, under rocks, and on low fo­liage in these areas. Some in­di­vid­u­als have been found around ponds or in deep bur­rows with a tur­ret of de­bris. (Jack­man, 1997; Milne and Milne, 1980; Parker, 1982)

Phys­i­cal De­scrip­tion

A typ­i­cal rabid wolf spi­der has a dark gray cephalotho­rax with two light lon­gi­tu­di­nal stripes ex­tend­ing across the top and a nar­row light line on each lat­eral mar­gin of the tho­rax. The ab­domen has a dark me­dian band, notched on each side in front of the mid­dle of the ab­domen, with sev­eral pairs of light spots on the rear part of the ab­domen. The first pair of legs is often black or dark brown, and the other legs are brown. Fe­males av­er­age 16 to 21 mm long, Males are typ­i­cally much smaller than fe­males, mea­sur­ing an av­er­age of 13 mm in length. (Com­stock and Gertsch, 1965; Milne and Milne, 1980)

  • Sexual Dimorphism
  • female larger
  • Average length
    males, 13 mm; females 21 mm
    in

De­vel­op­ment

Fe­male R. ra­bida drag their egg mass around, hav­ing spun a silken co­coon around the egg mass and at­tached it to spin­nerets. The co­coon dark­ens from shiny white to dirty brown be­fore the eggs hatch.

Spi­der­lings ride on their mother's back until they are ready for dis­per­sal. Egg sacs are from 7 to 10 mm in di­am­e­ter and con­tain from 168 to 365 eggs. (Jack­man, 1997; Milne and Milne, 1980)

Re­pro­duc­tion

Fe­male R. ra­bida re­lease a dragline. Males stum­ble across the line, which leads them to the fe­male. Once court­ing is over, the male spins a ball of silk and then re­leases sperm onto the ball. The fe­male at­taches the ball and drags it along with her until she is fer­til­ized. (Rovner, 1991)

Re­pro­duc­tion in R. ra­bida is one of the most com­monly de­bated and stud­ied areas of the an­i­mal's be­hav­ior. Rabid wolf spi­ders ef­fi­ciently de­tect move­ments. When males and fe­males are in courtship mode, they use a se­ries of dis­plays to in­ter­act and breed. R. ra­bida per­form a se­ries in­ter-bout turns in which one spi­der turns in a given di­rec­tion based upon the di­rec­tion in which the other spi­der turned pre­vi­ously. Pheromones also play a role in these in­tri­cate court­ing rit­u­als. Sci­en­tists have shown that the in­ten­sity of the pheromone is di­rectly re­lated to the turn­ing in males, specif­i­cally the de­gree and time of turn­ing.

Fe­male R. ra­bida drag their egg mass around, hav­ing spun a silken co­coon around the egg mass and at­tached it to spin­nerets. The co­coon dark­ens from shiny white to dirty brown be­fore the eggs hatch.

Spi­der­lings ride on their mother's back until they are ready for dis­per­sal. Egg sacs are from 7 to 10 mm in di­am­e­ter and con­tain from 168 to 365 eggs. (Jack­man, 1997; Milne and Milne, 1980; Rovner, 1991)

  • Breeding season
    Breeding occurs from late summer to early fall.
  • Range number of offspring
    168 to 365

The spi­der­lings hatch and re­main on the mother's back until they are ready for dis­per­sal. (Jack­man, 1997)

  • Parental Investment
  • precocial
  • pre-fertilization
    • provisioning
    • protecting
      • female
  • pre-hatching/birth
    • provisioning
      • female
    • protecting
      • female
  • pre-weaning/fledging
  • pre-independence
    • protecting
      • female

Lifes­pan/Longevity

Most rabid wolf spi­ders live up to around two years, six months of which is some­times spent on the mother's back. The larger, more fit R. ra­bida can live be­yond two years in sta­ble en­vi­ron­ments. (Com­stock and Gertsch, 1965)

  • Range lifespan
    Status: wild
    2 (high) years
  • Typical lifespan
    Status: wild
    2 (high) years

Be­hav­ior

Be­cause rabid wolf spi­ders are among the most com­mon spi­ders in the United States, they have adapted to many types of en­vi­ron­ments. These spi­ders are very ag­gres­sive to­wards hu­mans, spi­ders,other an­i­mals and in­sects. Be­cause of their wan­der­ing habits, these spi­ders are sub­ject to a plethora of in­ter­ac­tions be­tween the same and other species. Al­though harm­less to hu­mans, its bite is often feared. (Milne and Milne, 1980)

Com­mu­ni­ca­tion and Per­cep­tion

Rabid wolf spi­ders com­mu­ni­cate in many dif­fer­ent ways. One way is through the re­lease of pheromones. Both males and fe­males lay out a dragline and de­posit a chem­i­cal at­trac­tor on the line. Male wolf spi­ders in­ter­sect these lines and use their palps to fol­low the line for mat­ing. An­other type of com­mu­ni­ca­tion is web vi­bra­tions. This type of com­mu­ni­ca­tion, known as the sub­stra­tum-cou­pled vi­bra­tion sys­tem, is used mainly by males to at­tract fe­males, but is also used for males to com­mu­ni­cate with one an­other. Es­sen­tially, a male "plucks" the web fibers to play a "song". (Com­stock and Gertsch, 1965; Milne and Milne, 1980; Rovner, 1991; Un­known, 1999)

Food Habits

Rabid wolf spi­ders usu­ally pre­fer to eat small in­sects and other in­ver­te­brates. They have been known to eat crick­ets, lo­custs, ants, grasshop­pers and even other spi­ders. R. ra­bida is very strong and is not usu­ally in­tim­i­dated by larger or­gan­isms. These spi­ders are "sit-and-wait" preda­tors. In order to more eas­ily de­tect vi­sual and vi­bra­tory cues from prey, R. ra­bida must re­main mo­tion­less. These spi­ders do not eat solid ma­te­r­ial, so once they over­take the in­sect, they usu­ally suck out liq­uids and nu­tri­ents of their prey. (Jack­man, 1997; Parker, 1982; Rovner, 1989)

  • Animal Foods
  • insects
  • terrestrial non-insect arthropods

Pre­da­tion

While Ra­bidosa ra­bida is no­madic and moves fre­quently, it does so in a man­ner that is in­con­spicious to larger preda­tors. By stay­ing low in the brush, under lit­ter or ground cover, the spi­der can move ef­fi­ciently and most often re­mains un­de­tected. Wolf spi­ders get their name from they way they stalk their prey slowly and from a dis­tance, which has been as­so­ci­ated with wolves and other wild dogs. (Parker, 1982; Rovner, 1989)

  • Known Predators
    • Other Wolf Spiders
    • Dogs
    • Cats
    • Snakes

Ecosys­tem Roles

Ra­bidosa ra­bida has a lim­ited role in th ecosys­tem. Its main job as a preda­tor is to con­trol the boom­ing in­sect pop­u­la­tion. How­ever, as an oc­ca­sional prey species, it may con­tribute to feed­ing other or­gan­isms also.

Eco­nomic Im­por­tance for Hu­mans: Pos­i­tive

Al­though it is of lit­tle di­rect ben­e­fit to hu­mans, R. ra­bida is very com­mon among semi-ur­ban and rural areas and is very ac­tive in its pre­da­tion of in­sects. This can be a big ben­e­fit where pests are a prob­lem.

  • Positive Impacts
  • controls pest population

Eco­nomic Im­por­tance for Hu­mans: Neg­a­tive

Ra­bidosa ra­bida has lit­tle ad­verse af­fect on hu­mans out­side of being pests. These spi­ders are not poi­so­nous but have been know to bite un­sus­pect­ing vic­tims.

  • Negative Impacts
  • household pest

Con­ser­va­tion Sta­tus

Rabid wolf spi­ders are not cur­rently en­dan­gered or at any pre­v­e­lant risk of be­com­ing threat­ened.

Con­trib­u­tors

Nancy Shef­ferly (ed­i­tor), An­i­mal Di­ver­sity Web.

Justin Scar­bor­ough (au­thor), South­west­ern Uni­ver­sity, Stephanie Fab­ri­tius (ed­i­tor), South­west­ern Uni­ver­sity.

Glossary

Nearctic

living in the Nearctic biogeographic province, the northern part of the New World. This includes Greenland, the Canadian Arctic islands, and all of the North American as far south as the highlands of central Mexico.

World Map

acoustic

uses sound to communicate

agricultural

living in landscapes dominated by human agriculture.

bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

carnivore

an animal that mainly eats meat

chemical

uses smells or other chemicals to communicate

delayed fertilization

a substantial delay (longer than the minimum time required for sperm to travel to the egg) takes place between copulation and fertilization, used to describe female sperm storage.

ectothermic

animals which must use heat acquired from the environment and behavioral adaptations to regulate body temperature

forest

forest biomes are dominated by trees, otherwise forest biomes can vary widely in amount of precipitation and seasonality.

insectivore

An animal that eats mainly insects or spiders.

motile

having the capacity to move from one place to another.

native range

the area in which the animal is naturally found, the region in which it is endemic.

nomadic

generally wanders from place to place, usually within a well-defined range.

oviparous

reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.

pheromones

chemicals released into air or water that are detected by and responded to by other animals of the same species

seasonal breeding

breeding is confined to a particular season

sexual

reproduction that includes combining the genetic contribution of two individuals, a male and a female

suburban

living in residential areas on the outskirts of large cities or towns.

tactile

uses touch to communicate

temperate

that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).

terrestrial

Living on the ground.

urban

living in cities and large towns, landscapes dominated by human structures and activity.

vibrations

movements of a hard surface that are produced by animals as signals to others

visual

uses sight to communicate

young precocial

young are relatively well-developed when born

Ref­er­ences

Com­stock, J., W. Gertsch. 1965. The Spi­der Book. Ithaca, New York: Com­stock Pub­lish­ing Com­pany, Inc..

Jack­man, J. 1997. A Field Guide to Com­mon Texas In­sects. Hous­ton: Gulf Pub­lish­ing Com­pany.

Milne, L., M. Milne. 1980. The Audubon So­ci­ety Field Guide to North Amer­i­can In­sects and Spi­ders. New York: Chan­ti­cleer Press, Inc..

Parker, S. 1982. Syn­op­sis and Clas­si­fi­ca­tion of Liv­ing Or­gan­isms Vol­ume 2. Mc­Graw-Hill Book Com­pany.

Rovner, J. 1991. Turn­ing be­hav­ior dur­ing pheromone-stim­u­lated courtship in wolf spi­ders. An­i­mal Be­hav­iour, 42: 1015-1016.

Rovner, J. 1989. Wolf Spi­ders lack mir­ror-im­age re­spon­sive­ness seen in jump­ing spi­ders. An­i­mal Be­hav­iour, 38: 526-533.

Un­known, 1999. "In­tro­duc­tion to Ethol­ogy" (On-line). Ac­cessed Oct. 3, 2001 at http://​cas.​bellarmine.​edu/​tietjen/​Ethology/​introduction_​to_​ethology.​htm.